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RNA signals are shaped by where transcripts are in their lifecycle 1 . In eukaryotic cells, RNAs are transcribed and processed in the nucleus, then exported to the cytoplasm by the nuclear pore complex 2 . Exported molecules include messenger RNAs (mRNAs), ribosomal RNAs (rRNAs), transfer RNAs (tRNAs), small nuclear RNAs (snRNAs), micro RNAs (miRNAs), miRNAs, long noncoding RNAs (lncRNAs) and…